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Once your final files are uploaded, the early version of your manuscript, if you opted to have an early version of your article, will be published online. The date of the early version will be your article's publication date. The final article will be published to the same URL, and all versions of the paper will be accessible to readers.

We hope to receive your revised manuscript within 60 days. If you anticipate any delay, please let us know the expected resubmission date by replying to this email. Please note that revising your submission and if it goes towards acceptance, there is no need to protect anonymity. Your revised manuscript will be shared online when it is accepted.

If you would like to make changes to your version of this file, please go to https://www.plosgenetics.org/article/info%3Adoi%2F10.1371%2Fjournal.pgen.1005947 . Make sure to include the full range of permission statements below:

If you would like to make changes to your version of the manuscript, please note those changes must be made before the peer reviews are completed. If you're willing to make those changes, please state so in your revision. Note that published peer reviews are not attached to our decision to publish, as the review process is complete before we send them to our editors.

If you would like to make changes to your version of the manuscript, please describe these changes in detail, and we will ensure that the criteria, including high-quality documentation, are met for resubmission.

If you would like to make changes to your version of this file, please go to Referees Comments -> Uploaded Files and make your changes. If you feel any files are being retained from previous versions, please upload these as separate files and delete them from your version.

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